Abstract:Food caching behavior of birds is influenced by many factors, of which, the type of food and season are the two most critical factors. In this study, we used artificial feeding to explore food selection and food-caching behavior by varied tit (Parus varius) on pine and sunflowers seeds and to reveal the effects of the type of food and seasonal changes on the microhabitat use and spatial distribution of food storage sites. The results showed that varied tit preferred to store pine nuts, but during spring, they also stored a small percentage of sunflower seeds. They preferred to store the food within 100 m from the feeder, in a scattered spatial distribution. The frequency of stored food within 100 m decreased with the increase in the distance between food storage sites and feeders. The microhabitat of food caching included five kinds of locations: bark fissures, shrub's root bark, grass, leaf blades, and moss. Both models of food caching and micro habitat selection in varied tits were likely to be influenced by the season. The seasonal variations in the food-caching model was probably related to the decline in the available pine trees in natural environments, while the increasing usage of bark fissures as food-caching habitats was possibly related to the snow cover on the land. In general, varied tits preferred to store food in the proximity of the feeder in order to increase the efficiency of food caching. However, individual variation in the spatial use of food caching and the effect of seasonal variations require further study.